phd courses university of copenhagen

• The PhD programme

PhD courses

Course program for the PhD School

Read the course program for the PhD School here: Course program for the PhD School (pdf)

Planned courses

Courses offered by the Phd school are announced in the planned courses web calendar .

Introductory course including the course Good Scientific Practice and Quality of Legal Research (mandatory for all PhDs)

The PhD school organizes a introductory course to all newly enrolled PhD students once in the beginning of every semester. Please read more here.

Pedagogical course (mandatory for all PhDs)

A career as a researcher is also a career as an educator. Therefore, a pedagogical course is a mandatory part of your doctoral programme. The course is titled ‘ Learning principles at the faculty’ or ‘ Læringsprincipper på fakultetet’ (in Danish)’. The course is organised by the Study Services and is held before semester start. For more information about the course, please contact the Study Services ( [email protected] ).

The Faculty of Law, together with the PhD School and individual research centres, offer high quality PhD courses with the aim of establishing an active research environment that meets international standards and contributes to an attractive educational environment for PhD students.

The amount of course activity in which the PhD student shall participate has been rated as approx. 30 ECTS points.

The PhD Committee approves PhD courses and ensures their quality and continuous evaluation.

The Danish Legal Research Education Programme (JurForsk cooperation)

The PhD School participates in the Danish Legal Research Education Programme which offers a wide variety of courses relevant for PhD students within the field of law. It also offers a database of all current PhD students enrolled at the Danish JurForsk institutions as well as a newsletter function. For further information, please visit www.jurforsk.dk .

JurforskNordic.org: This website (hosted by the Danish Legal Research Education Programme, JurForsk) offers a platform for announcing PhD courses, conferences and seminars to Nordic PhD students in law. It is supported by a large number of Nordic institutions and offers a newsletter function. For further information, please visit www.jurforsknordic.org .

Generic courses

The PhD student may sign up for generic courses offered by the PhD school at the Faculty of Law or by any other PhD School at the University of Copenhagen. Further information about generic courses can be found at the web pages of each PhD School.

ECTS points for other courses and conferences

PhD students can apply for ECTS points for attending other courses and conferences according to the guidelines adopted by the PhD Committee. Enrolled PhD students can find these guidelines on the intranet.

To apply for ECTS points please use this application form .

PhD roundtables

Obligatory PhD roundtable seminars are conducted within the framework of the individual research centers at the faculty. They aim to better integrate PhD students in the activities of the faculty’s research centers. Additionally, the PhD roundtables provide PhD students with an opportunity to present their research projects to colleagues who possess professional expertise in the specific area of law as well as to other academic scholars who wish to participate in the roundtable seminars.

The roundtables are mandatory for PhD students. The PhD roundtables should run for at least one hour. The presentation of the PhD project takes place within the first six months of PhD studies and satisfies the obligation under §4.4.3 of the PhD Regulations.

Following their participation in a PhD roundtable seminar, the PhD student should include information regarding such participation in their first status report after the seminar. Additionally, subsequent to their presentation at a PhD roundtable, the PhD student must submit a one-page report to the head of the PhD School elaborating to what extent s/he considers the feedback they received from other participants as being valuable for the purposes of his/her continued research activities.

The schedule of PhD roundtables that will take place in individual research centers is available on the webpages of the PhD School and the relevant research centers.

The roundtable seminars are planned by the PhD student and are announced at  https://jura.ku.dk/phd/

Presenting your research project at the roundtable seminar grants 1 ECTS point.

Language courses

The University provides students with the opportunity to undertake language courses, mainly in Danish and English

The courses are offered by Studieskolen and the Centre for Internationalisation and Parallel Language Use ( CIP ).

Planned courses

Open up your academic publishing toolbox, phd seminar on qualitative research methods in legal studies, become a more productive and joyful writer: workshop on ph.d. writing, phd summer school 2024.

Show all events

Graduate School of Health and Medical Sciences

Key numbers from the Graduate School

Browse the PhD Course Catalogue

Join our Introduction Course

Choose between 22 graduate programmes

Promoting Responsible Conduct of Research

Apply for enrolment.

Thesis and defence

For PhD supervisors

Talk for phd students and phd supervisors: seeing positive in negative results, prolonged case handling at the graduate school, talk for students and supervisors: seeing positive in negative results, symposium on metabolic bone disease (deadline 17 march).

Show all news

PhD defences

Phd defence: stefanie alexandra eberle, phd defence: anna pouplier, phd defence: julie kjems.

Show all events

Department of Sociology

• PhD studies

PhD courses

The PhD programme includes coursework of approximately one semester of study, totaling 30 ECTS.

There are “generic” (general) courses arranged at Graduate School of Social Sciences . The topics of these courses are relevant to all of the faculty's PhD students. The students are to gain general competence in, e.g. pedagogy, communications, writing scientific papers in English, etc. The sociology courses, on the other hand, are typically offered by the department. See the website: Research – PhD student – PhD courses. They are usually first announced by e-mail and also posted at www.phdcourses.dk.

Taking a generic introductory course and a sociological project design course in the first semester of the  program is mandatory. The department regularly offers courses in sociological theory and methods (sometimes in cooperation with the Sociology departments at Aalborg and Lund University).

PhD students may also take courses at other departments and universities, both nationally ( https://phdcourses.dk/ ) and internationally.

Further, PhD students are encouraged to participate in the suggestion and planning of course activities. This ensures that the specialised courses offered resonate with the interests of the PhD students, while providing the opportunity for the students to become part of an international academic network and build their capacity to organise academic events. On KUnet under the department you can find a description on how to plan a PhD course.

Ph.D.-courses

How to plan a course (pdf in Danish).

• Faculty of Theology
• The PhD programme

PhD courses

Phd courses offered by or in collaboration with the faculty of theology:.

• 6 - 7 February 2024 Supervision of Bachelor Projects and Theses Registration deadline: 19 January 2024 Link: Supervision of Bachelor Projects and Theses
• 8 – 19 April 2024 Grundlæggende universitetspædagogik for ph.d.-studerende (in Danish) Registration deadline: 28 February 2024 Link: Grundlæggende universitetspædagogik for ph.d.-studerende
• 2 May 2024 Open up your Academic Publishing Toolbox Registration deadline: 24 April 2024 Link:   Open up your Academic Publishing Toolbox
• 29 May 2024 Philosophy and mysticism between past and present Registration deadline: 8 May 2024 Link:  Philosophy and mysticism between past and present
• 13 June 2024 Open up your Academic Publishing Toolbox Registration deadline: 5 June 2024 Link:   Open up your Academic Publishing Toolbox
• 17 - 18 December 2024 Vejledning af bachelorprojekter og specialer Registration deadline: 13 November 2024 Link: Vejledning af bachelorprojekter og specialer

PhD courses in Denmark:

• PhD course offered by the PhD school at The Faculty of Humanities
• The national portal ( phdcourses.dk)  allows you to search for PhD courses offered by all Danish universities.

PhD School in the Humanities

The PhD School in the Humanties

Intranet for phd students and staff, how to obtain a phd scholarship, phd courses, phd thesis, assessment and defence, international applicants and international visiting phd scholars.

• Department of Economics
• PhD Programme

PhD courses

The PhD students take half a year’s worth of PhD courses beyond the master's degree requirement. These courses provide specialised knowledge about the dissertation topic as well as generic competencies within research, teaching and knowledge dissemination. The courses can be of shorter or longer duration, and can be offered in direct connection with the programme, by the Graduate School, by national and international graduate schools, or by other relevant higher education institutions.

For some years, the large majority of PhD courses have been offered by the two national graduate schools within the fields of Economics and Finance. These two schools are characterized by offering courses of high academic standard, covering broader foundational courses taught by the best Danish researchers as well as narrower topics courses drawing on leading Danish and international researchers. Each school offers many courses in its respective field to nicely meet the PhD students’ course demand.

• PhD courses offered at the Department of Economics
• Copenhagen Graduate School of Social Sciences

Department of Biology

• BIO - English
• PhD programme

  PhD Programme

Scholarships

Phd study at the department of biology.

The Department of Biology is among the largest Departments at the University of Copenhagen. It employs 475 academic and technical staff and hosts research programmes and teaching curricula that cover all major biological sub-disciplines. University rankings typically place the field of biology/biochemistry at the University of Copenhagen first in Denmark and among the top 30 universities internationally.

The mission of the PhD programme at the Department of Biology is to offer world-class research training and education through participation in exciting, frontier research projects.

The successful candidates will join the lively group of 150 PhD students studying at the Department of Biology and participate in the many exciting events and activities organised by the PhD student network and the Department, including the annual PhD day, the biannual BIO conference, weekly research seminars, and career fairs.

All PhD students at Department of Biology are enrolled at the  PhD school of Science .

Local PhD coordinators Robin Andersson Tel: 3533 0245 ​ [email protected] Bioinformatic & RNA Biology A​nders Priemé​ Tel: 3533 0147 [email protected] Microbiology,  Cell-and Neurobiology,  Ecology and Evolution ​ Henriette Pilegaard Tel:  3532 1687 [email protected] Cell Biology and Physiology,  Genomics and Molecular Biomedicin Niels Daugbjerg  Tel: 3533 1830 [email protected] Freshwater Biology,  Marine  B​​iology,  Terrestrial Ecology​​​​ Michael Lisby Tel: 3532 2120 [email protected] Functional genomics Sine Lo Svenningsen Tel: 3532 2033 [email protected] Biomolecular  Sciences

PhD secretaries Jannike Dyrskjøt   (Molecular Biology and Genes, Molecular Cell Biology and Physiology, Protein Sciences) Jeannette Melsen   (Aquatic Sciences, Ecology and Evolution, Microbiology)

Department of Nutrition, Exercise and Sports

• PhD education

PhD courses

Upcomming PhD courses at Department of Nutrition, Exercise and Sports

PhD course: Introduction to Nutritional Metabolomics

Phd course: vascular function and angiogenesis in relation to health and physical activity, phd course: culture, sport and society, phd school of science, phd courses at the faculty of science, previous phd courses.

• Department of Anthropology
• PhD Programme

The PhD programme in Anthropology 

The Department of Anthropology at the University of Copenhagen takes a broad approach to social and cultural anthropology, reflected in the courses offered and in ongoing research activities. It provides diversity in terms of regional ethnographic concentration, theoretical orientations and methodologies. 

Department of Computer Science

• Department of Computer Science DIKU
• Research School

PhD Courses

Course Program

In addition to general purpose courses, such as Scientific Writing and Introduction to University Pedagogy, the IMLRS will strive to host PhD courses representing the research fields of the Image Section and IMLRS. Furthermore, the IMLRS aims to host an annual summer school with international guest lecturers on a specialized topic within imaging and machine learning.

• Course archive

Upcoming Courses

• International PhD course in Nonlinear Statistics

Copenhagen Center for Social Data Science (SODAS)

New phd position at sodas, phd position at the copenhagen center for social data science (sodas), university of copenhagen, with a focus on investigating the role of adults’ curiosity for decisions, behavior, and life outcomes, apply here: https://employment.ku.dk/phd/show=161571.

Who are we looking for? The Copenhagen Center for Social Data Science (SODAS; https://sodas.ku.dk/ ), Faculty of Social Sciences, University of Copenhagen, invites applications for a PhD position (100%, 36 months). Supported by the Tryg Foundation, the chosen candidate will primarily work in a project related to curiosity in children and adults, together with a PhD student at the Department of Psychology, a post-doc (at SODAS), Guido Makransky (Department of Psychology), and Ingo Zettler (SODAS and Department of Psychology).

The position is available from 1 July 2024, but a sooner or later starting date is also possible.

The chosen candidate will be part of the research project “Shedding New Light on Curiosity in Children and Adults”. The main goal of this project is to generate new knowledge that can support children and adults in having a positive life both mentally and socially, by increasing the understanding of: 1) how to best conceptualize and measure curiosity in children and adults; 2) how interventions can foster individual’s curiosity levels; and 3) the role of (different aspects of) curiosity for important outcomes. The project has three intertwined work packages focusing on psychometrics; Virtual Reality (VR) measurement and interventions with children; as well as survey, experimental, and register data studies with adults.

The chosen candidate will be primarily responsible for organizing and conducting the studies with adults. The chosen candidate is expected to publish scientific articles in international journals as well as to present findings at national and international conferences. In general, the position is research-focused, and funding is available for conducting studies, conference attendances, and a research visit such that the chosen candidate can start building their international network and research expertise. For the chosen candidate, there will be possibilities to influence the project and develop new project ideas within the project frame. In addition, the chosen candidate will also be asked to fulfill other work obligations (typically teaching or supervising thesis students) to a small degree in the overall period.

Applicants should have a master degree in computational social science, social data science,  psychology, another social science discipline, or adjacent disciplines. Next to interest in conducting research on curiosity, applicants should have experience with conducting survey and/or experimental studies, including quantitative analyses. Having experience with register data is an additional strength, but no precondition. It is expected that applicants will publish in internationally recognized scientific journals in English, and that the chosen candidate will contribute continuously with their collaboration skills to the interdisciplinary, international research team. High proficiency in spoken and written English is thus a requirement.

What do we offer? The chosen candidate will become part of the SODAS research environment and active doctoral program with about 20 PhD students. SODAS is an interdisciplinary research, teaching, and impact center, combining traditional social science theories and methods with novel data science methods and paradigms. In collaboration with partners in Denmark and abroad, SODAS is home to research projects funded by, among others, the European Research Council (ERC) as well as both independent and private Danish research foundations. SODAS hosts the English-language Social Data Science (SDS) Master degree program at the Faculty of Social Sciences.

The working language at SODAS is English. Our community maintains strong collaborative relationships with all Departments at the Faculty of the Social Sciences (Anthropology, Economics, Political Science, Psychology, Sociology), the Department of Computer Science at the University of Copenhagen, as well as internationally renowned, computational social science research groups and centers in the Copenhagen area, including the Networks, Data, and Society (NERDS) group at the IT University of Copenhagen, the Social Complexity Lab at the Technical University of Denmark, and the Pioneer Centre for AI which operates across all Danish universities.

How to apply Submit a complete application at our online portal. Click on the “Apply now” icon at the bottom of the page to apply. The documents must be in Adobe PDF or Word.

• Cover/motivation letter (max. 3 pages)
• Curriculum vitae (including publications and conference presentations, if applicable)
• Diplomas and transcript of grades (bachelor and master, if applicable)
• An abstract of the master thesis is written in English (1 page)
• Master thesis (if applicable)
• Up to 3 research articles and/or working papers (if applicable)

Please note that it is only possible to upload one document per attachment category. If more than one document has to be uploaded in the same category, please make sure that they are scanned and collected into one file.  

Terms of Employment

• In order to be awarded a PhD scholarship the chosen candidates have to enroll as PhD students at the Faculty of Social Sciences, cf. the rules of the Danish Ministerial order No 1039 of 27 August 2013. Most likely, the applicants will be enrolled in the PhD program at the Department of Psychology, University of Copenhagen.   
• Further information about the PhD study programme is available on the website of Copenhagen Graduate School of Social Sciences:   https://samf.ku.dk/phd-skolen/english/  
• Under "Legal basis" on the website you will find information about the rules and guidelines for the PhD programme, and the Danish Ministerial Order on the PhD Programme at the Universities.    
• The salary range starts at approximately DKK 29,000 per month + a 17.1 % contribution to the pension scheme. The exact net salary depends on various aspects including the chosen pension scheme and taxation but can be expected to be around DKK 21,000 net/month. It is possible to negotiate salary supplements on an annual basis.

The Recruitment Process Shortlisted applicants are notified of the composition of the assessment committee, and each applicant has the opportunity to comment on the part of the assessment that relates to the applicant themselves. Further information on these recruitment process at University of Copenhagen can be found here:   https://employment.ku.dk/faculty/recruitment-process/

An Equal Opportunity Workplace The University of Copenhagen is committed in its pursuit of academic excellence to equality of opportunity and to creating an inclusive working environment and therefore encourages all qualified candidates to apply, regardless of personal background, gender, sexual orientation, age, disability, ethnicity etc. For more on the diverse working place environment at the University and the University’s participation in the HRS4R HR Excellence in Research, see https://employment.ku.dk/working-at-ucph/eu-charter-for-researchers/

International Applicants The University of Copenhagen offers information and services for international researchers. Please find more information about these services as well as information on entering and working in Denmark here: https://ism.ku.dk/

Contact Information Information about the recruitment process is available from Dorthea Andersen, HR, e-mail: [email protected] , please refer to ID number: 211-1797/24-2H #1 .

Additional information about the position can be obtained from Ingo Zettler, e-mail: [email protected] The closing date for applications is 30 April 2024 at 23:59   CEST.

Applications or material received after the deadline will not be taken into account.

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• Published: 01 April 2024

Complexity of avian evolution revealed by family-level genomes

• Josefin Stiller   ORCID: orcid.org/0000-0001-6009-9581 1 ,
• Shaohong Feng   ORCID: orcid.org/0000-0002-2462-7348 2 , 3 , 4 , 5 ,
• Al-Aabid Chowdhury 6 ,
• Iker Rivas-González   ORCID: orcid.org/0000-0002-0515-0628 7 ,
• David A. Duchêne   ORCID: orcid.org/0000-0002-5479-1974 8 ,
• Qi Fang   ORCID: orcid.org/0000-0002-9181-8689 9 ,
• Yuan Deng 9 ,
• Alexey Kozlov   ORCID: orcid.org/0000-0001-7394-2718 10 ,
• Alexandros Stamatakis   ORCID: orcid.org/0000-0003-0353-0691 10 , 11 , 12 ,
• Santiago Claramunt   ORCID: orcid.org/0000-0002-8926-5974 13 , 14 ,
• Jacqueline M. T. Nguyen   ORCID: orcid.org/0000-0002-3076-0006 15 , 16 ,
• Simon Y. W. Ho   ORCID: orcid.org/0000-0002-0361-2307 6 ,
• Brant C. Faircloth   ORCID: orcid.org/0000-0002-1943-0217 17 ,
• Julia Haag   ORCID: orcid.org/0000-0002-7493-3917 10 ,
• Peter Houde   ORCID: orcid.org/0000-0003-4541-5974 18 ,
• Joel Cracraft   ORCID: orcid.org/0000-0001-7587-8342 19 ,
• Metin Balaban 20 ,
• Uyen Mai 21 ,
• Guangji Chen   ORCID: orcid.org/0000-0002-9441-1155 9 , 22 ,
• Rongsheng Gao 9 , 22 ,
• Chengran Zhou   ORCID: orcid.org/0000-0002-9468-5973 9 ,
• Yulong Xie 2 ,
• Zijian Huang 2 ,
• Zhen Cao 23 ,
• Zhi Yan   ORCID: orcid.org/0000-0003-2433-5553 23 ,
• Huw A. Ogilvie   ORCID: orcid.org/0000-0003-1589-6885 23 ,
• Luay Nakhleh   ORCID: orcid.org/0000-0003-3288-6769 23 ,
• Bent Lindow   ORCID: orcid.org/0000-0002-1864-4221 24 ,
• Benoit Morel 10 , 11 ,
• Jon Fjeldså   ORCID: orcid.org/0000-0003-0790-3600 24 ,
• Peter A. Hosner   ORCID: orcid.org/0000-0001-7499-6224 24 , 25 ,
• Rute R. da Fonseca   ORCID: orcid.org/0000-0002-2805-4698 25 ,
• Bent Petersen   ORCID: orcid.org/0000-0002-2472-8317 8 , 26 ,
• Joseph A. Tobias   ORCID: orcid.org/0000-0003-2429-6179 27 ,
• Tamás Székely   ORCID: orcid.org/0000-0003-2093-0056 28 , 29 ,
• Jonathan David Kennedy 30 ,
• Andrew Hart Reeve   ORCID: orcid.org/0000-0001-5233-6030 24 ,
• Andras Liker 31 , 32 ,
• Martin Stervander   ORCID: orcid.org/0000-0002-6139-7828 33 ,
• Agostinho Antunes   ORCID: orcid.org/0000-0002-1328-1732 34 , 35 ,
• Dieter Thomas Tietze   ORCID: orcid.org/0000-0001-6868-227X 36 ,
• Mads Bertelsen 37 ,
• Fumin Lei   ORCID: orcid.org/0000-0001-9920-8167 38 , 39 ,
• Carsten Rahbek   ORCID: orcid.org/0000-0003-4585-0300 25 , 30 , 40 , 41 ,
• Gary R. Graves   ORCID: orcid.org/0000-0003-1406-5246 30 , 42 ,
• Mikkel H. Schierup   ORCID: orcid.org/0000-0002-5028-1790 7 ,
• Tandy Warnow 43 ,
• Edward L. Braun   ORCID: orcid.org/0000-0003-1643-5212 44 ,
• M. Thomas P. Gilbert   ORCID: orcid.org/0000-0002-5805-7195 8 , 45 ,
• Erich D. Jarvis 46 , 47 ,
• Siavash Mirarab   ORCID: orcid.org/0000-0001-5410-1518 48 &
• Guojie Zhang   ORCID: orcid.org/0000-0001-6860-1521 2 , 3 , 5 , 49  

Nature ( 2024 ) Cite this article

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We are providing an unedited version of this manuscript to give early access to its findings. Before final publication, the manuscript will undergo further editing. Please note there may be errors present which affect the content, and all legal disclaimers apply.

• Evolutionary biology
• Genome evolution
• Molecular evolution
• Phylogenetics

Despite tremendous efforts in the past decades, relationships among main avian lineages remain heavily debated without a clear resolution. Discrepancies have been attributed to diversity of species sampled, phylogenetic method, and the choice of genomic regions 1–3 . Here, we address these issues by analyzing genomes of 363 bird species 4 (218 taxonomic families, 92% of total). Using intergenic regions and coalescent methods, we present a well-supported tree but also a remarkable degree of discordance. The tree confirms that Neoaves experienced rapid radiation at or near the Cretaceous–Paleogene (K–Pg) boundary. Sufficient loci rather than extensive taxon sampling were more effective in resolving difficult nodes. Remaining recalcitrant nodes involve species that challenge modeling due to extreme GC content, variable substitution rates, incomplete lineage sorting, or complex evolutionary events such as ancient hybridization. Assessment of the impacts of different genomic partitions showed high heterogeneity across the genome. We discovered sharp increases in effective population size, substitution rates, and relative brain size following the K–Pg extinction event, supporting the hypothesis that emerging ecological opportunities catalyzed the diversification of modern birds. The resulting phylogenetic estimate offers novel insights into the rapid radiation of modern birds and provides a taxon-rich backbone tree for future comparative studies.

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Authors and affiliations.

Section for Ecology and Evolution, Department of Biology, University of Copenhagen, Copenhagen, Denmark

Josefin Stiller

Center for Evolutionary & Organismal Biology, & Women’s Hospital, Zhejiang University School of Medicine, Hangzhou, China

Shaohong Feng, Yulong Xie, Zijian Huang & Guojie Zhang

Liangzhu Laboratory, Zhejiang University Medical Center, Hangzhou, China

Shaohong Feng & Guojie Zhang

Department of General Surgery, Sir Run-Run Shaw Hospital, Zhejiang University School of Medicine, Hangzhou, China

Shaohong Feng

Innovation Center of Yangtze River Delta, Zhejiang University, Jiashan, China

School of Life and Environmental Sciences, University of Sydney, Sydney, New South Wales, Australia

Al-Aabid Chowdhury & Simon Y. W. Ho

Bioinformatics Research Centre, Aarhus University, Aarhus, Denmark

Iker Rivas-González & Mikkel H. Schierup

Center for Evolutionary Hologenomics, The Globe Institute, University of Copenhagen, Copenhagen, Denmark

David A. Duchêne, Bent Petersen & M. Thomas P. Gilbert

BGI-Shenzhen, Beishan Industrial Zone, Shenzhen, China

Qi Fang, Yuan Deng, Guangji Chen, Rongsheng Gao & Chengran Zhou

Computational Molecular Evolution Group, Heidelberg Institute for Theoretical Studies, Heidelberg, Germany

Alexey Kozlov, Alexandros Stamatakis, Julia Haag & Benoit Morel

Institute of Computer Science, Foundation for Research and Technology Hellas, Heraklion, Greece

Alexandros Stamatakis & Benoit Morel

Institute for Theoretical Informatics, Karlsruhe Institute of Technology, Karlsruhe, Germany

Alexandros Stamatakis

Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, Canada

Santiago Claramunt

Department of Natural History, Royal Ontario Museum, Toronto, Ontario, Canada

College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia

Jacqueline M. T. Nguyen

Research Institute, Australian Museum, Sydney, New South Wales, Australia

Department of Biological Sciences and Museum of Natural Science, Louisiana State University, Baton Rouge, LA, USA

Brant C. Faircloth

Department of Biology, New Mexico State University, Las Cruces, NM, USA

Peter Houde

Department of Ornithology, American Museum of Natural History, New York, NY, USA

Joel Cracraft

Bioinformatics and Systems Biology Graduate Program, University of California San Diego, La Jolla, CA, USA

Metin Balaban

Computer Science and Engineering, University of California San Diego, La Jolla, CA, USA

College of Life Sciences, University of Chinese Academy of Sciences, Beijing, China

Guangji Chen & Rongsheng Gao

Department of Computer Science, Rice University, Houston, TX, USA

Zhen Cao, Zhi Yan, Huw A. Ogilvie & Luay Nakhleh

Natural History Museum Denmark, University of Copenhagen, Copenhagen, Denmark

Bent Lindow, Jon Fjeldså, Peter A. Hosner & Andrew Hart Reeve

Center for Global Mountain Biodiversity, Globe Institute, University of Copenhagen, Copenhagen, Denmark

Peter A. Hosner, Rute R. da Fonseca & Carsten Rahbek

Centre of Excellence for Omics-Driven Computational Biodiscovery (COMBio), Faculty of Applied Sciences, AIMST University, Bedong, Kedah, Malaysia

Bent Petersen

Department of Life Sciences, Imperial College London, Silwood Park, Ascot, UK

Joseph A. Tobias

Milner Centre for Evolution, University of Bath, Bath, UK

Tamás Székely

ELKH-DE Reproductive Strategies Research Group, University of Debrecen, Debrecen, Hungary

Center for Macroecology, Evolution, and Climate, The Globe Institute, University of Copenhagen, Copenhagen, Denmark

Jonathan David Kennedy, Carsten Rahbek & Gary R. Graves

HUN-REN-PE Evolutionary Ecology Research Group, University of Pannonia, Veszprém, Hungary

Andras Liker

Behavioural Ecology Research Group, Center for Natural Sciences, University of Pannonia, Veszprém, Hungary

Bird Group, Natural History Museum, Akeman St, Tring, Hertfordshire, United Kingdom

Martin Stervander

CIIMAR/CIMAR, Interdisciplinary Centre of Marine and Environmental Research, University of Porto, Porto, Portugal

Agostinho Antunes

Department of Biology, Faculty of Sciences, University of Porto, Porto, Portugal

NABU, Berlin, Germany

Dieter Thomas Tietze

Centre for Zoo and Wild Animal Health, Copenhagen Zoo, Frederiksberg, Denmark

Mads Bertelsen

Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

College of Life Science, University of Chinese Academy of Sciences, Beijing, China

Institute of Ecology, Peking University, Beijing, China

Carsten Rahbek

Danish Institute for Advanced Study, University of Southern Denmark, Odense, Denmark

Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA

Gary R. Graves

University of Illinois Urbana-Champaign, Champaign, IL, USA

Tandy Warnow

Department of Biology, University of Florida, Gainesville, FL, USA

Edward L. Braun

University Museum, NTNU, Trondheim, Norway

M. Thomas P. Gilbert

Vertebrate Genome Lab, The Rockefeller University, New York, NY, USA

Erich D. Jarvis

Howard Hughes Medical Institute, Durham, NC, USA

University of California, San Diego, San Diego, CA, USA

Siavash Mirarab

Villum Center for Biodiversity Genomics, Department of Biology, University of Copenhagen, Copenhagen, Denmark

Guojie Zhang

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Corresponding authors

Correspondence to Josefin Stiller , Siavash Mirarab or Guojie Zhang .

Supplementary information

Supplementary information.

This file contains Supplementary Methods and Supplementary Results.

Reporting Summary

Peer review file, supplementary data.

Table of all sequenced species with taxonomic grouping according to Howard & Moore. 4th Edition and accession numbers of the used genome assemblies. Given as a separate tab-delimited text file.

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Stiller, J., Feng, S., Chowdhury, AA. et al. Complexity of avian evolution revealed by family-level genomes. Nature (2024). https://doi.org/10.1038/s41586-024-07323-1

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Published : 01 April 2024

DOI : https://doi.org/10.1038/s41586-024-07323-1

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• Department of Psychology

The PhD programme

The PhD programme at the Department of Psychology is a three year full-time research programme fully or partially funded by external funds or by the Department.

If you wish to apply for a PhD scholarship please read the information about the PhD programme on the following pages:

It is not a requirement that you speak Danish in order to enrol as a PhD student af the Department of Psychology.

About the PhD programme

Application for a phd scholarship, phd programme regulations, phd courses, the organisation of the phd programme, graduate school.

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U.S. News & World Report Ranks UH Graduate Programs Among Nation’s Best in 2024

Four UH Graduate Programs Rank in Top 10, 17 in Top 50

By Shawn Lindsey — 713-743-5725

• University and Campus
• Student, Faculty and Staff Success

The University of Houston is once again proving that it is a powerhouse in graduate education, with 13 of its programs securing spots among the top 50 nationwide in the 2024 U.S. News & World Report graduate program rankings, released today. Three engineering programs maintain their top 50 rank pending the release of 2024 engineering rankings, which have been delayed. In total, UH has 17 programs in the top 50.

A total of four UH graduate programs rank in the top 10 nationwide, including three from the UH Law Center: Health Care Law (No. 7), Part-Time Law (No. 8) and Intellectual Property Law (No. 8). The Petroleum Engineering program in the Cullen College of Engineering maintains its No. 9 ranking pending forthcoming rankings in the discipline. Since 2020, UH has seen a 70% increase in the number of graduate programs earning a top 50 ranking.

“Being acknowledged among the nation's top performers in these vital fields is truly gratifying and affirms the national prestige of these programs and the caliber of faculty, staff and students,” said Diane Z. Chase, UH senior vice president for academic affairs and provost.  “Delivering top-tier professional and graduate programs and positioning students for success in their respective fields of graduate study is integral to our vision of ascending as a top 50 public university.”

This year, seven UH Law Center specialty programs are ranked among the Top 50 in the country, with three ranked No. 1 in the state of Texas.

Top 50 Ranked UH Graduate Programs Nationwide:

• Health Care Law (No. 7)
• Part-Time Law (No. 8)
• Intellectual Property Law (No. 8)
• Petroleum Engineering (No. 9)*
• Trial Advocacy (No. 13)
• Online Overall Graduate Education Program (No. 14)
• Legal Writing (No. 16)
• Environmental Law (No. 30)
• Tax Law (No. 32)
• Chemical Engineering (No. 34)*
• International Law (No. 38)
• Pharmacy (No. 41)
• Dispute Resolution Law (No. 42)
• Business Part-Time MBA (No. 45)
• Speech-Language Pathology (No. 44)
• Industrial Manufacturing (No. 50)*
• Political Science (No. 50)**

*2023 ranking; U.S. News has delayed 2024 engineering program rankings

**discipline not ranked in 2024 but maintains a previous top 50 ranking

Each year, U.S. News ranks professional school programs in business, education, engineering, law, medicine, and nursing, including specialties in each area. The rankings are based on enrollment numbers, job placement rates, faculty statistics, and other essential quality indicators designed to help prospective students make informed decisions.

“Deciding where to attend graduate school can be formidable, considering the wealth of schools with distinctive and quality programs,” said LaMont Jones, Ed.D., managing editor for Education at U.S. News. “The Best Graduate Schools rankings and related content empower prospective students to choose the best fi t for their educational needs and ultimately achieve career success.”

The top programs in the nation earn “best of” recognition by U.S. News. UH made the following 2024 graduate program lists:

• Best Business Schools
• Best Health Schools
• Best Public Affairs Schools
• Best Science Schools
• Best Law Schools
• Best Education Schools

To see all of the graduate program rankings, or to learn more about the ranking methodology, visit the  U.S. News & World Report website .

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April 09, 2024

March 29, 2024

Three Renowned Argonne Scientists Accept Joint Appointments at the University of Houston

Three top researchers with the U.S. Department of Energy’s Argonne National Laboratory have accepted joint appointments in various capacities at the University of Houston. This strategic collaboration furthers critical research efforts, public-private partnerships and educational opportunities for students.

• Science, Energy and Innovation

March 28, 2024

Predicting Weight Gain in Children Using Second Generation Anti-Psychotic Medication

Research from the University of Houston College of Pharmacy indicates that preventing clinically significant weight gain in children using second-generation antipsychotic medicine may be possible.

• Health and Medicine

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• The Integrated PhD Pro...

The Integrated  MSc and  PhD programme (3+5)

The Integrated MSc and PhD programme makes it possible to integrate the MSc programmes’ optional programme elements and the writing of the PhD dissertation. This allows students and their supervisors more time for the research element of the PhD programme.

The former 4+4 PhD is now a part of the Integrated PhD Scheme (3+5).

Admission requirements

The Integrated PhD programme is open to applicants who:

• have completed the Bachelors degree with a good result.
• are enrolled on an MSc programme offered by SCIENCE  or  have been offered admission to an MSc programme offered by SCIENCE
• can coordinate and integrate the remaining ECTS of their MSc programme with the proposed PhD programme.
• is qualified to be admitted to the PhD programme
• have not  started writing the MSc thesis.

The academic content of the MSc programme must be related to the proposed PhD programme.  Read more requirements in the programme description below. 

Organisation of the Integrated MSc/PhD programme

The prescribed period of study for the integrated programme is five full-time equivalents (FTEs) (two for the MSc programme and three for the PhD programme). If, at commencement of the integrated 3+5 scheme, the student has passed one or more programme elements, the period of study is reduced correspondingly.

The 3+5 scheme is divided into two parts:

Part A: The student completes the MSc programme and concurrently completes one third of the PhD programme (one FTE). Part A is completed when the student completes the MSc programme and must be organized in such a way that the MSc programme is completed two years before completion of the entire integrated programme of study.

Part B : The student is a PhD student exclusively and completes the PhD programme in accordance with the approved PhD plan. Part B can be commenced once the student has completed the MSc programme (part A), and the regular assessments have been satisfactory, meaning that the student has lived up to the requirements in the PhD plan.

You can find more information here.

Find information on financing here

USC retains #1 international MBA ranking, climbs in nursing, education

U.s. news rankings show broad-based excellence in graduate programs.

For the 11th straight year, the University of South Carolina has earned the No. 1 ranking in the country for its International MBA program, according to the Best Graduate School rankings released April 9 by U.S. News & World Report. This year marks the 35th consecutive year that the program has ranked in the top three nationally.

“The Darla Moore School of Business’ No. 1-ranked International MBA offers students an interdisciplinary and comprehensive curriculum that prepares them to lead multinational organizations,” said Rohit Verma, the Moore School’s dean. “Our IMBA program is led by the top thought leaders in the international business discipline and delivers high job placement rates and competitive salaries with global corporations.”

The U.S. News graduate program rankings show broad-based excellence at the University of South Carolina, with additional business programs also ranked and significant jumps in nursing and education.

At the Darla Moore School of Business, the graduate-level operations and supply chain program ranked No. 16, up four spots from last year. The Moore School is also ranked No. 3 by Gartner for its undergraduate program. In addition, the professional MBA program is at the No. 19 spot in the latest U.S. News rankings.

The College of Nursing continued a climb it has seen over the past few years, earning the No. 22 spot for its overall master’s program, up from No. 32 last year. The College of Nursing was also ranked No. 1 in January for its online master’s program — for the fourth straight year. The college’s students perform in the top 1 percent nationally on the NCLEX, the national licensure exam for registered nurses.

“We are proud of the work being done by our faculty and staff to create an array of nationally recognized graduate programs, which provide superior education for our students who will become tomorrow’s academic and industry leaders."

The College of Education ranked No. 29 in the category of Best Education Schools, with its graduate programs moving up 11 spots from No. 40 last year. The college has improved by 58 spots in the rankings over the past two years.

In other rankings highlights, the Joseph F. Rice School of Law ranks No. 13 for trial advocacy, up from No. 15 last year and No. 30 for its clinical training programs.

“Graduate education is deeply important to the University of South Carolina, and our continued recognition in U.S. News rankings reflects the university’s culture of academic excellence,” says Provost Donna Arnett. “We are proud of the work being done by our faculty and staff to create an array of nationally recognized graduate programs, which provide superior education for our students who will become tomorrow’s academic and industry leaders.”

The University of South Carolina’s ranked graduate and professional programs include diverse areas of study in the sciences, humanities, technology, engineering, health sciences, law and business. These programs help train graduate students destined to become future leaders in their fields, providing the skilled workforce needed to propel the state’s economy.

USC has more nationally ranked graduate programs than all other colleges and universities in the state combined.

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Uri college of nursing graduate programs ranked among nation’s best.

Doctor of Nursing Practice ranks 66 nationally; Master’s program remains in top 50

KINGSTON, R.I. — April 9, 2024 — The University of Rhode Island College of Nursing graduate programs are again ranked among the top of nursing colleges in the country, making an impressive year-over-year rise, according to the latest U.S. News & World Report rankings.

The College’s Doctor of Nursing Practice program is tied at number 66 of 354 nursing schools across the country with a DNP program, up from number 86 last year, according to the rankings released April 9. The College’s master’s program also continued its impressive performance in recent years. For the third year in a row, the program ranks among the top 50 in the nation, slotting in at number 48 among 603 nursing schools surveyed.

“We are committed to providing an excellent education to all our students, including our graduate students who benefit from our top-notch facilities and some of the best educators in their field,” said Dean Danny Willis . “Education is transformative, and students are not just a number to us. We get to know them and facilitate their professional and personal success. It is very rewarding to see the College’s efforts recognized on a national level.”

U.S. News & World Report , among the leading authorities in college and university rankings, surveyed 648 nursing schools with master’s or doctoral programs accredited by either the Accreditation Commission for Education in Nursing or the Commission on Collegiate Nursing Education. The publication determined each school’s overall rank using 15 distinct ranking factors, including faculty credentials, student-to-faculty ratio, faculty research prowess and nursing practice experience, assessment scores from peer institutions, and more.

The URI College of Nursing met and exceeded the standards as it continues to increase its offerings at the undergraduate and graduate level. About one-third of the URI College of Nursing’s faculty members are recognized Fellows in national nursing organizations, including the American Academy of Nursing, for their distinguished academic achievements. Dozens of URI students were inducted in the Sigma Theta Tau honor society for nursing last year, and a handful were inducted into the Honor Society of Phi Kappa Phi. The number of faculty publications, funded external grants and committed gifts to the College have all increased over the last five years as the College continues to have major impacts on health and health care.

The advancements the College has made are obvious not only in the graduate programs, but in its undergraduate offerings as well. The College’s B.S. in Nursing program tied at number 67 out of 656 programs in the country for the second year in a row, in rankings released Sept. 17. The College has made an impressive rise in the rankings, climbing from number 96 just two years ago, the first year the publication ranked undergraduate programs, to 67 last year. Visit the URI College of Nursing Academics website for more information on all educational offerings.

University of South Florida

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Nearly two dozen graduate programs at USF considered among the best in America

• April 9, 2024

University News

By Althea Johnson , University Communications and Marketing

The University of South Florida is home to nearly two dozen graduate programs considered among the best in America, according to new rankings released today by U.S. News & World Report. USF features 23 graduate programs ranked inside the top 100 among all public and private institutions, including 11 ranked in the top 50.

USF’s highest-ranked programs are industrial and organizational psychology at No. 3, criminology at No. 18 and audiology, which comes in at No. 22. 

Among USF Health’s ranked programs, nursing anesthesia jumped 58 spots into the top 50, the physical therapy program rose by double digits to No. 33 and the nursing master’s program now sits in the top 25 at No. 24. 

In addition, USF’s social work and part-time MBA programs both saw double-digit gains and the education program broke into the top 50.  

“We are thrilled that many University of South Florida graduate programs are included among the nation’s best in U.S. News & World Report’s rankings,” USF President Rhea Law said. “This recognition underscores the commitment of our faculty and staff, who continue to provide high-quality programs that empower our students to enhance their skills, expand their knowledge and progress in their careers by earning an advanced degree.”

Full list of USF’s top 50 programs:

Industrial and Organizational Psychology* – 3 Criminology* – 18 Audiology – 22 Nursing – Master’s – 24 Library and Information Studies* – 27 Public Health – 29 Physical Therapy – 33 Nursing – Doctor of Nursing Practice – 34 Nursing – Anesthesia – 36 Rehabilitation Counseling* – 46 Education – Overall – 48

Full list of USF’s 51-100 programs:

Healthcare Management*– 52 Speech-Language Pathology – 52 Business ‐ Part‐time MBA – 53 Earth Sciences* – 54 Pharmacy – 60 Physician's Assistant/Associate Studies* – 65 Fine Arts* – 73 Social Work – 83  Statistics* – 86  Business – Full-time MBA – 87  Sociology* – 92 Psychology* – 97  

As U.S. News does not release new rankings for every program, every year, rankings denoted with an asterisk have been republished this year. U.S News also announced that medical, engineering and clinical psychology school rankings will be released at a later date. 

According to U.S. News, rankings are generated through “ranking indicators” derived from reputation and/or data reported by each institution.  For some programs, U.S. News evaluates key areas such as expert opinions in the field, opportunities for student engagement, resources for student support and career success, as well as faculty and student performance, in terms of existing credentials, research impact and metrics such as student job placement.

In addition to the graduate program rankings, in September 2023 U.S. News ranked USF as one of the nation’s top 50 public universities for the fifth consecutive year , and the university earned its highest ranking ever among all universities public or private. 

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USF in the News

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March 22, 2024

More USF in the News

U.S. News ranks 16 graduate programs, specialties at IU among nation’s top 10, 32 among top 20

Excellence abounds through Indiana University. And once again, this excellence has deservedly been recognized nationally.

Today, IU graduate programs have been ranked among the best in the country in the 2024-25 U.S. News & World Report Best Graduate Schools rankings . Demonstrating IU’s academic and research impact and leadership, IU leads the state in the total number of top 10 and top 20 graduate programs or specialties ranked by U.S. News.  

Only seven public universities nationally, and only two schools in the Big Ten, have more top 20 programs and specialty areas than IU.

Preparing the next generation of global public affairs leaders

IU’s acclaimed  O’Neill School of Public and Environmental Affairs  leads all U.S. public affairs graduate schools with the most No. 1-ranked specialty areas. 

The O’Neill School has the top-ranked programs in environmental policy and management, public finance and budgeting, public management and leadership, and nonprofit management. It also boasts the top two rankings in the nonprofit management category, with the school’s program on the IU Bloomington campus ranking No. 1 and its program on the IU Indianapolis campus ranking No. 2.   

Thirteen specialty areas at the O’Neill School ranked in the top 20, with nine Bloomington specialties placing in the top 10 and four Indianapolis specialties ranking in the top 20.

The O’Neill School’s master’s program in public affairs ranks No. 1 among public institutions and No. 2 among all universities.

National excellence recognized across IU 

In total, 16 IU graduate programs and specialties rank among the nation’s top 10, with 32 programs and specialties ranking among the top 20. Both figures represent increases from last year and make IU the state’s leader for the total number of top 10 and top 20 program and specialties ranked.  

The  Kelley School of Business  ranked fifth among public institutions and 20th among all colleges and universities, the school’s highest ranking since 2012. The school had two specialties ranked among the top 10, entrepreneurship and accounting, and seven specialties in the top 20. 

The  School of Education  ranks in the top 20 in seven of eight ranked specialties, including four in the top 10: student counseling and personnel services, elementary teacher education, secondary teacher education, and higher education administration. 

Finally, the  College of Arts and Sciences, McKinney School of Law , and  Luddy School of Informatics, Computing and Engineering  each have top 20-ranked specialty programs. 

These rankings provide much deserved national recognition of IU’s world-class faculty and staff, who lead our progress and transformation, empowering our students to reach their fullest potential, serve the public good, and advance our society.

Pamela Whitten

President Indiana University

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