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A doctorate is the pinnacle of an arts and science education. Founded in 1886, the Graduate School of Arts and Science at NYU is among the oldest schools offering doctoral programs in the United States. Today NYU’s doctoral programs span the humanities, sciences, and social sciences, and students pursue cutting-edge research with the close supervision of NYU’s internationally recognized research faculty. New York City resources complement and enhance our vibrant intellectual communities. Use the links below to explore Doctor of Philosophy and dual advanced degrees at New York University.

Ph.D. Programs Dual Degree Programs

Graduate Course Search

Use this tool to search for courses by semester. Click on a course in the resulting list to go to a page with more information about the course. Syllabi are also available here and on each course page. Note that syllabi are provided for planning purposes only and are subject to change without notice. The course instructor will provide an updated syllabus to all registered students. Every effort is made to keep this information accurate and current, but students should consult  Albert  for official offering information.

BIOL-GA 1001 Bio Core I: Molecular Systems | syllabus

BIOL-GA 1002 Bio Core II | syllabus

BIOL-GA 1004 Environmental Health

BIOL-GA 1005 Ecotoxicology

BIOL-GA 1006 Toxicology

BIOL-GA 1007 Programming for Biologists | syllabus

BIOL-GA 1009 Biological Databases and Data Mining | syllabus

BIOL-GA 1011 Immunobiology | syllabus

BIOL-GA 1020 Advanced Topics in Cellular and Molecular Immunology

BIOL-GA 1023 Hot Topics in Infectious Diseases | syllabus

BIOL-GA 1031 Special Topics in Physiology and Disease | syllabus

BIOL-GA 1032 Metabolic Disorders | syllabus

BIOL-GA 1045 Protein Biochemistry | syllabus

BIOL-GA 1051 Cell Biology: The Nucleus and Beyond | syllabus

BIOL-GA 1052 Frontiers in Microbiology | syllabus

BIOL-GA 1069 Principles of Evolution | syllabus

BIOL-GA 1072 Molecular Controls of Organismal Form and Function

BIOL-GA 1080 Emerging Pathogens | syllabus

BIOL-GA 1082 Genes and Animal Behavior | syllabus

BIOL-GA 1122 Molecular Lab Biology I | syllabus

BIOL-GA 1123 Molecular Lab Biology II | syllabus

BIOL-GA 1124 Molecular Lab Biology III

BIOL-GA 1125 Molecular Lab Biology IV

BIOL-GA 1127 Bioinformatics and Genomes | syllabus

BIOL-GA 1128 Systems Biology | syllabus

BIOL-GA 1129 Evolutionary Genetics and Genomics | syllabus

BIOL-GA 1130 Applied Genomics: Introduction to Bioinformatics and Network Modeling | syllabus

BIOL-GA 1131 Biophysical Modeling of Cells and Populations | syllabus

BIOL-GA 1132 Genomics of Human Populations | syllabus

BIOL-GA 1140 BIOL- GA 1140 Genomic Innovation | syllabus

BIOL-GA 2003 Bio Core III: Molecules and Cells | syllabus

BIOL-GA 2004 Bio Core IV | syllabus

BIOL-GA 2005 Current Topics in Biology I: Precision Medicine

BIOL-GA 2006 Current Topics in Biology II: Gatekeepers of Cell Division and Cancer

BIOL-GA 2007 Current Topics in Biology III: Evolution and Medicine

BIOL-GA 2008 Current Topics in Biology IV | syllabus

BIOL-GA 2015 Genomics and Global Public Health | syllabus

BIOL-GA 2021 Biomaterials - Tissue Interface II

BIOL-GA 2024 Biomaterials - Tissue Interface I

BIOL-GA 2026 Introduction to Electron Microscopy

BIOL-GA 2029 Metal and Ceramic Biomaterials

BIOL-GA 2030 Statistics in Biology | syllabus

BIOL-GA 2031 Statistics and Machine Learning for Genomics

BIOL-GA 2040 Molecular and Genetic Toxicology

BIOL-GA 2048 Integrative Seminars in Oral Biology I: Oral Systemic Connections

BIOL-GA 2049 Integrative Seminars in Oral Biology II: Bone Biology and Craniofacial Development

BIOL-GA 2130 Developmental and Stem Cell Systems I

BIOL-GA 2303 Intro to Biostatistics

BIOL-GA 2310 Principles of Toxicology

BIOL-GA 3001 Art of Scientific Investigation | syllabus

BIOL-GA 3015 Pre-Doctoral Colloquium

BIOL-GA 3015 Pre-Doctoral Colloquium Srvy Lab Mthd

BIOL-GA 3034 Predoctoral Colloquium: Laboratory Rotation

BIOL-GA 3303 Research

BIOL-GA 3304 Research

BIOL-GA 3305 Reading

BIOL-GA 3306 Reading

BIOL-GA 3340 Reading in Oral Biology

BIOL-GA 3350 Directed Individual Research in Oral Biology I

BIOL-GA 3351 Directed Individual Research in Oral Biology II

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Biology PhD Program

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NYU Shanghai, in partnership with the NYU Graduate School of Arts and Science and the NYU Department of Biology, invites applications from exceptional students for PhD study and research in Biology.   Participating students are enrolled in the NYU GSAS Biology PhD program, complete their coursework at the NYU Department of Biology in New York, and then transition to full-time residence at NYU Shanghai where they undertake their doctoral research under the supervision of NYU Shanghai faculty.

Highlights of the Program

• NYU degree upon graduation
• Graduate coursework at the NYU Department of Biology in New York
• Research opportunities with and close mentorship by NYU Shanghai faculty
• Access to the vast intellectual resources of NYU GSAS and NYU Department of Biology
• Cutting-edge research environment at NYU Shanghai, including a thriving community of PhD students, post-doctoral fellows, and research associates, activities such as a regular program of seminars and visiting academics, and links with other universities within and outside China
• Financial aid through the NYU Shanghai Doctoral Fellowship , including tuition, fees, and an annual stipend
• Additional benefits exclusive to the NYU Shanghai program, including international health insurance, housing assistance in New York, and travel funds

Genomics, Bioinformatics, Evolutionary Analysis of Large-Scale Genomics Data, Molecular Evolution, Computational Biology

Jungseog Kang

Chromosome Segregation, Mitosis, Antimitotic Cancer Drug Screen

Recent Publications by NYU Shanghai Faculty

• KBase: The United States Department of Energy Systems Biology Knowledgebase.Nature biotechnology 36 (7), 566, 2018
• Pangolin genomes and the evolution of mammalian scales and immunity Genome research 26(10):1312-1322, 2016
• Unknown unknowns: essential genes in quest for function. A Danchin, G Fang, Microbial biotechnology 9 (5), 530-540 2016
• Comparative analysis of the transcriptome across distant species Nature 512 (7515), 445 166 2014
• Transcriptomic and phylogenetic analysis of a bacterial cell cycle reveals strong associations between gene co-expression and evolution G Fang, KD Passalacqua, J Hocking, PM Llopis, M Gerstein, NH Bergman, BMC genomics 14 (1), 450, 2014

Jungseog Kang

• Improving drug discovery with high-content phenotypic screens by systematic selection of reporter cell lines. Kang J et. al. Nature Biotechnology (2016) 34:70-77.
• Mitotic centromeric targeting of HP1 and its binding to Sgo1 are dispensable for sister-chromatid cohesion in human cells. Kang J et. al. Mol. Biol. Cell (2011) 22:1181-1190.
• Kinase signaling in the spindle checkpoint. Kang J et. al. J. Biol. Chem. (2009) 284:15359-63.

Selected Faculty and Student Features

" Communicating Science: NYU Shanghai’s First Postdoctoral and Doctoral Research Assembly " (Xiaoai Lyu)

" A New Way to Screen Cancer Drugs " (Jungseog Kang)

" Professor Gang Fang Wins NYU GIPH Research Affinity Group Network Challenge " (Gang Fang)

Structure of Program

Participating students complete the PhD degree requirements set by the NYU Department of Biology and in accordance with the academic policies of NYU GSAS. Each student develops an individualized course and research plan in consultation with the Director of Graduate Study at the NYU Department of Biology and the student’s NYU Shanghai faculty advisor. A typical sequence follows:

Begin program with funded research rotation, up to 3 months preceding first Fall semester, to familiarize with NYU Shanghai and faculty as well as lay a foundation for future doctoral study.

Complete PhD coursework at Department of Biology alongside other NYU PhD students. 

Return to Shanghai for second funded research rotation to solidify relationships with NYU Shanghai faculty and make further progress in research.

Under supervision of NYU Shanghai faculty advisor, pursue dissertation research and continue coursework. Depending on each student’s individualized course of study, return visits to New York may also occur. Complete all required examinations and progress evaluations, both oral and written, leading up to submission and defense of doctoral thesis.

To learn more about the NYU Biology PhD program degree requirements, please visit this page .

Application Process and Dates

Applications are to be submitted through the NYU GSAS Application portal , within which students should select the Biology PhD as their program of interest, and then indicate their preference for NYU Shanghai by marking the appropriate checkbox when prompted. Applicants will be evaluated by a joint admissions committee of New York and Shanghai faculty. Application requirements are set by the NYU Department of Biology and are the same as those for all NYU PhD applicants, no matter their campus preference; however, candidates are recommended to elaborate in their application and personal statements about their specific interests in the NYU Shanghai program and faculty.

For admission in Fall 2024, the application deadline is December 1, 2023.

Interested students are welcome to contact Vivien Du , PhD Program Manager, at [email protected] with any inquiries or to request more information.

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Application Process

Potential Biology Fellowship candidates apply using the standard GSAS online application for admission to the Biology PhD program. There is no separate application for the NYUAD Global PhD Student Fellowship.

Interested students complete a laboratory rotation in Abu Dhabi during their first year of study, and Fellowship recipients will be notified by the end of their first year in the PhD program.

Visit the  GSAS Application Resource Center for more information on how to apply.

The application deadline for Fall 2024 is December 1, 2023. Please note that applications are only accepted for the fall semester, as we do not offer Spring admission.

For more information about our programs, please contact  [email protected] .

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Properly Write Your Degree

The correct way to communicate your degree to employers and others is by using the following formats:

Degree - This is the academic degree you are receiving. Your major is in addition to the degree; it can be added to the phrase or written separately.  Include the full name of your degree, major(s), minor(s), emphases, and certificates on your resume.

Double Majors - You will not be receiving two bachelor's degrees if you double major. Your primary major determines the degree (Bachelor of Arts or Bachelor of Science). If you're not fully sure which of your majors is primary, check CheckMarq or call the registrar's office.

Example: Primary Major: Psychology ; Secondary Major: Marketing

• Bachelor of Arts Degree in Psychology & Marketing

Primary Major: Marketing ; Secondary Major: Psychology

• Bachelor of Science Degree in Marketing & Psychology

In a letter, you may shorten your degree by writing it this way:

• In May 20XX, I will graduate with my Bachelor's degree in International Affairs.
• In December 20XX, I will graduate with my Master's degree in Counseling Education.

Not sure which degree you are graduating with? Here is a list of Undergraduate Majors and corresponding degrees:

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NYU Sends Out Offers of Admission To the Class of 2028

Offers of admission to NYC campus were made to just 8% of the 118,000 undergraduate applicants

NYU has released offers of admission to the Class of 2028. Offers of admission to NYU’s New York campus were made to just 8% of those submitting 118,000 applications to be members of next fall’s entering undergraduate class. Three of NYU’s undergraduate colleges offered admission to fewer than 5% of applicants. 

This cycle, NYU witnessed its largest early decision applicant pool in history, comprising over 22,000 applications, marking a 56%-surge in Early Decision applications across NYU’s degree-granting campuses over the past five years. Over 60% of the applications to NYU were from public schools, with almost 9,000 of them from students who work with non-profit organizations such as OppNet, Bottom Line, College Advising Corps (CAC), Minds Matter, and the Charles Hayden Foundation, among others, who partner with NYU to create new pathways to increase educational access.   

NYU anticipates that 20% of the class will be Pell Grant recipients and 20% will be First-Generation students. This year’s cohort represents over 103 countries, 49 states (all but North Dakota), several US territories, and approximately 1,000 students from New York City public schools.

The NYU Promise: The NYU Promise is President Linda G. Mills’ expanded commitment to affordability. More than 1,000 students are recipients of the NYU Promise, which offers free tuition for students with family incomes under $100,000. In addition, the NYU Promise includes meeting students’ demonstrated need if it is beyond tuition and increasing aid annually to keep pace with increases in tuition.

“NYU’s admitted class of 2028 is made up of students who represent the best of us,” said Jonathan Williams, Associate Vice President of Undergraduate Admissions, Pre-College Access and Pathways. “Throughout their high school journey, these remarkable young individuals hail from various corners of the United States and the globe. They have proven themselves to be exemplary bridgebuilders, striving to connect diverse perspectives and foster understanding. Their unwavering perseverance in the face of myriad challenges showcases their dedication to creating a better world – together. They are shining examples of good listeners, global citizens, and future leaders. Their presence will undoubtedly enrich the NYU community and contribute to a brighter future for all.”

New York University

Founded in 1831, NYU is one of the world’s foremost research universities (with more than $1 billion per year in research expenditures, it is ranked seventh among private research universities) and is a member of the selective Association of American Universities. NYU has degree-granting university campuses in New York, Abu Dhabi, and Shanghai; has 13 other global academic sites, including London, Paris, Florence, Tel Aviv, Buenos Aires, and Accra, and US sites in Washington, DC, Los Angeles, CA, and Tulsa, OK; and both sends more students to study abroad and educates more international students than any other U.S. college or university. Through its numerous schools and colleges, NYU is a leader in conducting research and providing education in the arts and sciences, law, medicine, business, dentistry, engineering, education, nursing, the cinematic and performing arts, music and studio arts, public administration, social work, public health, and professional studies, among other areas.

Press Contact

Marine Biology

• College of the Environment
• University of Washington

SAFS Undergraduate Community Building Event May 1st

The School of Aquatic and Fishery Sciences DEI committee will be hosting an event on May 1, 2024, from 3:30 to 5:00 pm  for undergraduates to build community with graduate students and postdocs in the SAFS department.

This informal event will have graduate students and postdocs stationed in the FSH building lobby with posters containing information on research opportunities, professional development, graduate school applications, and much more. Undergrads are invited to walk around and introduce themselves to the grads and postdocs and gain insight on any topics of interest. Light snacks and refreshments will be provided.

Please see the attached flyer and  RSVP here

We look forward to seeing you there! 🙂

New Director of the Florence S. Downs PhD Program in Nursing Research & Theory Development

April 11, 2024.